$0.55, AJo, top pair facing overbet shove on the turn

PokerStars, $0.49 + $0.06 - Hold'em No Limit - 3,500/7,000 (625 ante) - 9 players

UTG: 284,186 (41 bb)
UTG+1: 84,160 (12 bb)
MP: 173,906 (25 bb) VPIP 30, PFR 10, 179 hands
MP+1: 629,613 (90 bb)
HJ: 136,780 (20 bb)
CO: 170,804 (24 bb)
BU (Hero): 254,077 (36 bb)
SB: 184,608 (26 bb)
BB: 259,770 (37 bb)

Pre-Flop: (16,125) Hero is BTN with J♥ A♠
2 players fold, MP calls 7,000, 3 players fold, Hero raises to 21,000, 2 players fold, MP calls 14,000

Flop: (58,125) 9♥ 2♦ A♦ (2 players)
MP checks, Hero bets 21,000, MP calls 21,000

Turn: (100,125) 4♥ (2 players)
MP bets 131,281 (all-in), BU (Hero) folds

Total pot: 100,125
MP wins 100,125

Hi,

I noticed that the villain had a tendency to overplay their pairs. However they were mostly calling, I didn't notice them playing aggressively before. I couldn't figure out what his bet meant - perhaps he wanted to protect against double flush draw out there? In general I found that calling big bets with single pair hands rarely works out at these stakes. Can you guys find a call here?

Thanks!